The Truck Number (or Truck Factor) is the number of people with key knowledge that you cannot replace i.e. if that number of people went simultaneously under a truck you wouldn't be able to carry on developing.
I believe that certain, German chemical companies forbid key members of staff from travelling together for this very reason...
Discussion here: http://c2.com/cgi/wiki?TruckNumber
Here's a story about Bill Atkinson being the one key person in the Mac's truck factor - one of the key people that worked on QuickDraw during the early days of the Mac. Had a car accident apparently and people were concerned that he wouldn't be able to finish his work on the Mac's graphics software:
http://folklore.org/StoryView.py?project=Macintosh&story=I_Still_Remember_Regions.txt
A high truck number is better - i.e. it's harder to wipe out that many critical people at once...
A low truck number is worse - i.e. there is a greater risk that a few critical people could be ill, or leave or die, leaving the project in a state of unrecoverable collapse.
Pair progamming is a good way to distribute critical knowledge around a team so that the truck number is as high as possible.