This is caused by a Trigger in the default template for TreeViewItem:
<MultiTrigger>
<MultiTrigger.Conditions>
<Condition Property="IsSelected" Value="true"/>
<Condition Property="IsSelectionActive" Value="false"/>
</MultiTrigger.Conditions>
<Setter Property="Background" TargetName="Bd" Value="{DynamicResource {x:Static SystemColors.ControlBrushKey}}"/>
<Setter Property="Foreground" Value="{DynamicResource {x:Static SystemColors.ControlTextBrushKey}}"/>
</MultiTrigger>
Removing this from the default Aero theme template leaves:
<PathGeometry x:Key="TreeArrow" Figures="M0,0 L0,6 L6,0 z"/>
<Style x:Key="ExpandCollapseToggleStyle" TargetType="{x:Type ToggleButton}">
<Setter Property="Focusable" Value="False"/>
<Setter Property="Width" Value="16"/>
<Setter Property="Height" Value="16"/>
<Setter Property="Template">
<Setter.Value>
<ControlTemplate TargetType="{x:Type ToggleButton}">
<Border Width="16" Height="16" Background="Transparent" Padding="5,5,5,5">
<Path x:Name="ExpandPath" Fill="Transparent" Stroke="#FF989898" Data="{StaticResource TreeArrow}">
<Path.RenderTransform>
<RotateTransform Angle="135" CenterX="3" CenterY="3"/>
</Path.RenderTransform>
</Path>
</Border>
<ControlTemplate.Triggers>
<Trigger Property="IsMouseOver" Value="True">
<Setter Property="Stroke" TargetName="ExpandPath" Value="#FF1BBBFA"/>
<Setter Property="Fill" TargetName="ExpandPath" Value="Transparent"/>
</Trigger>
<Trigger Property="IsChecked" Value="True">
<Setter Property="RenderTransform" TargetName="ExpandPath">
<Setter.Value>
<RotateTransform Angle="180" CenterX="3" CenterY="3"/>
</Setter.Value>
</Setter>
<Setter Property="Fill" TargetName="ExpandPath" Value="#FF595959"/>
<Setter Property="Stroke" TargetName="ExpandPath" Value="#FF262626"/>
</Trigger>
</ControlTemplate.Triggers>
</ControlTemplate>
</Setter.Value>
</Setter>
</Style>
<ControlTemplate x:Key="MyTemplate" TargetType="{x:Type TreeViewItem}">
<Grid>
<Grid.ColumnDefinitions>
<ColumnDefinition MinWidth="19" Width="Auto"/>
<ColumnDefinition Width="Auto"/>
<ColumnDefinition Width="*"/>
</Grid.ColumnDefinitions>
<Grid.RowDefinitions>
<RowDefinition Height="Auto"/>
<RowDefinition/>
</Grid.RowDefinitions>
<ToggleButton x:Name="Expander" Style="{StaticResource ExpandCollapseToggleStyle}" ClickMode="Press" IsChecked="{Binding IsExpanded, RelativeSource={RelativeSource TemplatedParent}}"/>
<Border x:Name="Bd" SnapsToDevicePixels="true" Grid.Column="1" Background="{TemplateBinding Background}" BorderBrush="{TemplateBinding BorderBrush}" BorderThickness="{TemplateBinding BorderThickness}" Padding="{TemplateBinding Padding}">
<ContentPresenter x:Name="PART_Header" HorizontalAlignment="{TemplateBinding HorizontalContentAlignment}" SnapsToDevicePixels="{TemplateBinding SnapsToDevicePixels}" ContentSource="Header"/>
</Border>
<ItemsPresenter x:Name="ItemsHost" Grid.Column="1" Grid.ColumnSpan="2" Grid.Row="1"/>
</Grid>
<ControlTemplate.Triggers>
<Trigger Property="IsExpanded" Value="false">
<Setter Property="Visibility" TargetName="ItemsHost" Value="Collapsed"/>
</Trigger>
<Trigger Property="HasItems" Value="false">
<Setter Property="Visibility" TargetName="Expander" Value="Hidden"/>
</Trigger>
<Trigger Property="IsSelected" Value="true">
<Setter Property="Background" TargetName="Bd" Value="{DynamicResource {x:Static SystemColors.HighlightBrushKey}}"/>
<Setter Property="Foreground" Value="{DynamicResource {x:Static SystemColors.HighlightTextBrushKey}}"/>
</Trigger>
<Trigger Property="IsEnabled" Value="false">
<Setter Property="Foreground" Value="{DynamicResource {x:Static SystemColors.GrayTextBrushKey}}"/>
</Trigger>
</ControlTemplate.Triggers>
</ControlTemplate>